Ken Nagrod said:
A one year old child could easily count the growth rings in todays wood products. That and the fact that there isn't much development of the heartwood adds to seasonal expansion and contraction of wood products. Second, if you look at the side view of the cut you make in the molding, whether it's crown, base, chair rail, whatever, you will see seasonal movement in that direction (face to back) which does affect a tight joint over time. The across the grain movement doesn't just occur in the width of the molding but also in its thickness where you're still going across the grain structure.
Wood does not shrink equally in all directions. The ratio of movement tangentially versus radially is about 2-to-1. The statement about growth rings sounds really good at first, until you realize that there is very little movement in this direction to begin with, and therefore, the spacing of the growth rings has no bearing.
If you can see a change in the thickness of the material, it's not a seasonal problem, nor is it even a matter of installing wood that is dripping wet. I used the WoodWeb calculator for southern red oak going from 12% down to 8%, and the radial change in thickness for a 3/4" board is 0.005". That's not a seasonal change; that's putting wet wood on the wall and allowing it to dry, and it is still not considered measurable as far as wood products go.
As I said earlier, there are reasons for coping, but wood movement is not one of them, especially for flat moulding with a zero spring angle (i.e. chair rail). Understanding why something is done is just as important as knowing how to do it. When we lose sight of the reasons behind a method, we also lose sight of when it becomes important for doing it. As Gary pointed out (he posted after I wrote this), it is "speed" that is the primary driving factor for coping, and to that, I do not disagree.